About the Authors

Kathleen Semple Delaney

Katy_Delaney@nps.gov

Current address: Santa Monica Mountains National Recreation Area, National Park Service, Thousand Oaks, California, United States of America

Affiliation Department of Ecology and Evolutionary Biology, University of California Los Angeles, Los Angeles, California, United States of America

Seth P. D. Riley

Affiliation Santa Monica Mountains National Recreation Area, National Park Service, Thousand Oaks, California, United States of America

Robert N. Fisher

Affiliation San Diego Field Station, U.S. Geological Survey, San Diego, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KSD SPDR RNF. Performed the experiments: KSD. Analyzed the data: KSD. Contributed reagents/materials/analysis tools: KSD SPDR. Wrote the paper: KSD SPDR.