About the Authors

Aurelia Santoro

aurelia.santoro@unibo.it

Affiliations Department of Experimental Pathology, University of Bologna, Bologna, Italy, CIG-Interdepartmental Center for Biophysics and Biocomplexity Studies, University of Bologna, Bologna, Italy

Valentina Balbi

Affiliation Department of Experimental Pathology, University of Bologna, Bologna, Italy

Elisa Balducci

Affiliation Department of Experimental Pathology, University of Bologna, Bologna, Italy

Chiara Pirazzini

Affiliation Department of Experimental Pathology, University of Bologna, Bologna, Italy

Francesca Rosini

Affiliation Department of Experimental Pathology, University of Bologna, Bologna, Italy

Francesca Tavano

Affiliation Department of Experimental Pathology, University of Bologna, Bologna, Italy

Alessandro Achilli

Affiliations Department of Genetics and Microbiology, University of Pavia, Pavia, Italy, Department of Cell and Environmental Biology, University of Perugia, Perugia, Italy

Paola Siviero

Affiliation National Council Research, Institute of Neuroscience, Padova, Italy

Nadia Minicuci

Affiliation National Council Research, Institute of Neuroscience, Padova, Italy

Elena Bellavista

Affiliations Department of Experimental Pathology, University of Bologna, Bologna, Italy, CIG-Interdepartmental Center for Biophysics and Biocomplexity Studies, University of Bologna, Bologna, Italy

Michele Mishto

Affiliations Department of Experimental Pathology, University of Bologna, Bologna, Italy, CIG-Interdepartmental Center for Biophysics and Biocomplexity Studies, University of Bologna, Bologna, Italy, Institute of Biochemistry, Medical Faculty Charité, Berlin, Germany

Stefano Salvioli

Affiliations Department of Experimental Pathology, University of Bologna, Bologna, Italy, CIG-Interdepartmental Center for Biophysics and Biocomplexity Studies, University of Bologna, Bologna, Italy

Francesca Marchegiani

Affiliation Italian National Research Center for Aging (I.N.R.C.A.), Ancona, Italy

Maurizio Cardelli

Affiliation Italian National Research Center for Aging (I.N.R.C.A.), Ancona, Italy

Fabiola Olivieri

Affiliations Italian National Research Center for Aging (I.N.R.C.A.), Ancona, Italy, Department of Molecular Pathology and Innovative Therapies, Polytechnic University of Marche, Ancona, Italy

Benedetta Nacmias

Affiliation Department of Neurological and Psychiatric Sciences, University of Florence, Florence, Italy

Andrea Maria Chiamenti

Affiliation Regional Center for Cerebral Aging, Valdagno, Vicenza, Italy

Luisa Benussi

Affiliation NeuroBioGen Lab-Memory Clinic, “Centro S.Giovanni di Dio-Fatebenefratelli”, Brescia, Italy

Roberta Ghidoni

Affiliations NeuroBioGen Lab-Memory Clinic, “Centro S.Giovanni di Dio-Fatebenefratelli”, Brescia, Italy, Proteomics Unit, “Centro S.Giovanni di Dio-Fatebenefratelli”, Brescia, Italy

Giuseppina Rose

Affiliation Department of Cell Biology, University of Calabria, Rende, Cosenza, Italy

Carlo Gabelli

Affiliation Regional Center for Cerebral Aging, Valdagno, Vicenza, Italy

Giuliano Binetti

Affiliation NeuroBioGen Lab-Memory Clinic, “Centro S.Giovanni di Dio-Fatebenefratelli”, Brescia, Italy

Sandro Sorbi

Affiliation Department of Neurological and Psychiatric Sciences, University of Florence, Florence, Italy

Gaetano Crepaldi

Affiliation National Council Research, Institute of Neuroscience, Padova, Italy

Giuseppe Passarino

Affiliation Department of Cell Biology, University of Calabria, Rende, Cosenza, Italy

Antonio Torroni

Affiliation Department of Genetics and Microbiology, University of Pavia, Pavia, Italy

Claudio Franceschi

Affiliations Department of Experimental Pathology, University of Bologna, Bologna, Italy, CIG-Interdepartmental Center for Biophysics and Biocomplexity Studies, University of Bologna, Bologna, Italy

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AS AT CF. Performed the experiments: AS EB CP FR FT. Analyzed the data: AS VB EB PS NM GR GP. Contributed reagents/materials/analysis tools: AS AA EB MM SS FM MC FO BN AMC LB RG CG GB SS GC. Wrote the paper: AS VB GR AT CF.