About the Authors

Chenguang Wang

Affiliation Department of Statistics, University of Florida, Gainesville, Florida, United States of America

Zhong Wang

Affiliation Center for Statistical Genetics, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Jiangtao Luo

Affiliation Center for Statistical Genetics, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Qin Li

Affiliation Department of Statistics, University of Florida, Gainesville, Florida, United States of America

Yao Li

Affiliation Department of Statistics, West Virginia University, Morgantown, West Virginia, United States of America

Kwangmi Ahn

Affiliation Center for Statistical Genetics, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Daniel R. Prows

Affiliation Department of Pediatrics, University of Cincinnati College of Medicine, Cincinnati, Ohio, United States of America

Rongling Wu

rwu@hes.hmc.psu.edu

Affiliation Center for Statistical Genetics, Pennsylvania State University, Hershey, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DP RW. Performed the experiments: DP. Analyzed the data: CW ZW JL QL YL KA. Contributed reagents/materials/analysis tools: CW ZW JL QL YL KA. Wrote the paper: RW.