About the Authors

Victoria Horner

vhorner@emory.edu

Affiliation Living Links, Yerkes National Primate Research Center, Emory University, Lawrenceville, Georgia, United States of America

Darby Proctor

Affiliation Living Links, Yerkes National Primate Research Center, Emory University, Lawrenceville, Georgia, United States of America

Kristin E. Bonnie

Affiliation Department of Psychology, Beloit College, Beloit, Wisconsin, United States of America

Andrew Whiten

Affiliation Centre for Social Learning and Cognitive Evolution, School of Psychology, University of St Andrews, Fife, Scotland, United Kingdom

Frans B. M. de Waal

Affiliation Living Links, Yerkes National Primate Research Center, Emory University, Lawrenceville, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VKH DP KEB AW FBMdW. Performed the experiments: VKH DP. Analyzed the data: VKH DP FBMdW. Wrote the paper: VKH AW FBMdW. Contributed to discussion: VKH DP KEB AW FBMdW. Supervised the entire project: FBMdW.