About the Authors

Daniel N. Frank

Current address: Division of Infectious Diseases, University of Colorado School of Medicine, Aurora, Colorado, United States of America

Affiliations Department of Molecular, Cellular, and Developmental Biology, University of Colorado at Boulder, Boulder, Colorado, United States of America, Mucosal and Vaccine Research Program Colorado, Aurora, Colorado, United States of America

Leah M. Feazel

Affiliation Department of Molecular, Cellular, and Developmental Biology, University of Colorado at Boulder, Boulder, Colorado, United States of America

Mary T. Bessesen

Affiliations Mucosal and Vaccine Research Program Colorado, Aurora, Colorado, United States of America, Department of Veterans Affairs Medical Center, Denver, Colorado, United States of America, University of Colorado School of Medicine, Aurora, Colorado, United States of America

Connie S. Price

Affiliations University of Colorado School of Medicine, Aurora, Colorado, United States of America, Division of Infectious Diseases, Denver Health Medical Center, Denver, Colorado, United States of America

Edward N. Janoff

Affiliations Mucosal and Vaccine Research Program Colorado, Aurora, Colorado, United States of America, Department of Veterans Affairs Medical Center, Denver, Colorado, United States of America, University of Colorado School of Medicine, Aurora, Colorado, United States of America

Norman R. Pace

norman.pace@colorado.edu

Affiliations Department of Molecular, Cellular, and Developmental Biology, University of Colorado at Boulder, Boulder, Colorado, United States of America, Mucosal and Vaccine Research Program Colorado, Aurora, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DF ENJ NP. Performed the experiments: DF LMF. Analyzed the data: DF LMF ENJ NP. Contributed reagents/materials/analysis tools: DF MTB CSP. Wrote the paper: DF ENJ NP.