About the Authors

Paul C. Cross

pcross@usgs.gov

Affiliation Northern Rocky Mountain Science Center, United States Geological Survey, Bozeman, Montana, United States of America

Dennis M. Heisey

Affiliation National Wildlife Health Center, United States Geological Survey, Madison, Wisconsin, United States of America

Brandon M. Scurlock

Affiliation Wyoming Game and Fish Department, Pinedale, Wyoming, United States of America

William H. Edwards

Affiliation Wyoming Game and Fish Department, Laramie, Wyoming, United States of America

Michael R. Ebinger

Affiliation Big Sky Institute, Montana State University, Bozeman, Montana, United States of America

Angela Brennan

Affiliations Northern Rocky Mountain Science Center, United States Geological Survey, Bozeman, Montana, United States of America, Department of Ecology, Montana State University, Bozeman, Montana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BMS WHE. Performed the experiments: BMS WHE AB. Analyzed the data: PC DMH. Contributed reagents/materials/analysis tools: PC BMS WHE MRE AB. Wrote the paper: PC DMH BMS WHE MRE AB.