About the Authors

I. Martin Sheldon

I.M.Sheldon@swansea.ac.uk

Affiliation Institute of Life Science, School of Medicine, Swansea University, Singleton Park, Swansea, United Kingdom

Andrew N. Rycroft

Affiliation Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Belgin Dogan

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Melanie Craven

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

John J. Bromfield

Affiliation Institute of Life Science, School of Medicine, Swansea University, Singleton Park, Swansea, United Kingdom

Alyssa Chandler

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Mark H. Roberts

Affiliation Institute of Life Science, School of Medicine, Swansea University, Singleton Park, Swansea, United Kingdom

Sian B. Price

Affiliation Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Robert O. Gilbert

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Kenneth W. Simpson

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Competing Interests

I.M.S., R.O.B. and A.N.R. have had direct, and K.W.S. indirect, research grant funding from a veterinary pharmaceutical company (Pfizer Animal Health). However, those funds are entirely independent of the present study. Furthermore, this does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors (http://www.plosone.org/static/policies.action#sharing.)

Author Contributions

Conceived and designed the experiments: IMS AR ROG KWS. Performed the experiments: IMS AR BD MC JJB AMC MHR SBP KWS. Analyzed the data: IMS AR BD JJB KWS. Contributed reagents/materials/analysis tools: IMS KWS. Wrote the paper: IMS KWS.