About the Authors

Keiko Fujikawa

fujikawa@med.hokudai.ac.jp

Affiliations Department of Pathology and Immunology, Hokkaido University Graduate School of Medicine, Sapporo, Japan, Faculty of Health Science, Hokkaido University, Sapporo, Japan, Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Takeshi Iwata

Affiliation National Institute of Sensory Organs, National Hospital Organization Tokyo Medical Center, Tokyo, Japan

Kaoru Inoue

Affiliation Faculty of Health Science, Hokkaido University, Sapporo, Japan

Masakazu Akahori

Affiliation National Institute of Sensory Organs, National Hospital Organization Tokyo Medical Center, Tokyo, Japan

Hanako Kadotani

Affiliation Department of Pathology and Immunology, Hokkaido University Graduate School of Medicine, Sapporo, Japan

Masahiro Fukaya

Affiliation Department of Anatomy, Hokkaido University Graduate School of Medicine, Sapporo, Japan

Masahiko Watanabe

Affiliation Department of Anatomy, Hokkaido University Graduate School of Medicine, Sapporo, Japan

Qing Chang

Affiliation Department of Ophthalmology and Visual Sciences, Washington University School of Medicine, St. Louis, Missouri, United States of America

Edward M. Barnett

Affiliation Department of Ophthalmology and Visual Sciences, Washington University School of Medicine, St. Louis, Missouri, United States of America

Wojciech Swat

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KF TI KI. Performed the experiments: KF TI MA HK MF QC. Analyzed the data: KF TI KI MA MF MW EMB WAS. Contributed reagents/materials/analysis tools: KF TI KI MW WAS. Wrote the paper: KF KI EMB WAS.