About the Authors

Ruth D. Ellis

ellisru@niaid.nih.gov

Affiliation Malaria Vaccine Development Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Laura B. Martin

Current address: Novartis Vaccines Institute for Global Health, Siena, Italy

Affiliation Malaria Vaccine Development Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Donna Shaffer

Affiliation Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Carole A. Long

Affiliation Laboratory of Malaria and Vector Research, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Kazutoyo Miura

Affiliation Malaria Vaccine Development Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Michael P. Fay

Affiliation Biostatistics Research Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

David L. Narum

Affiliation Malaria Vaccine Development Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Daming Zhu

Affiliation Malaria Vaccine Development Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Gregory E. D. Mullen

Current address: The Rayne Institute, Division of Imaging Sciences, School of Medicine, King's College London, United Kingdom

Affiliation Malaria Vaccine Development Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Siddhartha Mahanty

Current address: Laboratory of Parasitic Diseases, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Bethesda, Maryland, United States of America

Affiliation Malaria Vaccine Development Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Louis H. Miller

Affiliation Malaria Vaccine Development Branch, National Institute of Allergy and Infectious Diseases/National Institutes of Health, Rockville, Maryland, United States of America

Anna P. Durbin

Affiliation Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Competing Interests

Dr. Laura Martin is currently an employee of Novartis Vaccines Institute. However Novartis played no role in this study, either as a funder or collaborator, and Dr. Martin's work on this project was completed before her affiliation with Novartis began. Neither this nor any other author's affiliation alters the adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LBM DS GEDM SM LHM APD. Performed the experiments: RDE LBM DS CAL KM DZ APD. Analyzed the data: RDE LBM DS KM MPF APD. Contributed reagents/materials/analysis tools: DLN. Wrote the paper: RDE KM MPF APD.