About the Authors

Kohei Yamada

Contributed equally to this work with: Kohei Yamada, Shinsuke Matsuzaki

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Shinsuke Matsuzaki

Contributed equally to this work with: Kohei Yamada, Shinsuke Matsuzaki

Affiliation United Graduate School of Child Development, Osaka University, Kanazawa University and Hamamatsu University School of Medicine, Suita, Osaka, Japan

Tsuyoshi Hattori

Affiliation Department of Molecular Neuropsychiatry, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Ryusuke Kuwahara

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Manabu Taniguchi

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Hitoshi Hashimoto

Affiliations United Graduate School of Child Development, Osaka University, Kanazawa University and Hamamatsu University School of Medicine, Suita, Osaka, Japan, Laboratory of Molecular Neuropharmacology, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan

Norihito Shintani

Affiliations Department of Molecular Neuropsychiatry, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Molecular Neuropharmacology, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan

Akemichi Baba

Affiliation Laboratory of Molecular Neuropharmacology, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan

Natsuko Kumamoto

Affiliation Department of Anatomy and Neuroscience, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Kazuo Yamada

Affiliation Laboratory of Molecular Psychiatry, RIKEN Brain Science Institute, Wako, Saitama, Japan

Takeo Yoshikawa

Affiliation Laboratory of Molecular Psychiatry, RIKEN Brain Science Institute, Wako, Saitama, Japan

Taiichi Katayama

katayama@ugscd.osaka-u.ac.jp

Affiliation United Graduate School of Child Development, Osaka University, Kanazawa University and Hamamatsu University School of Medicine, Suita, Osaka, Japan

Masaya Tohyama

Affiliations Department of Anatomy and Neuroscience, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, United Graduate School of Child Development, Osaka University, Kanazawa University and Hamamatsu University School of Medicine, Suita, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KY SM TK MT. Performed the experiments: KY TH RK MT KY. Analyzed the data: KY SM TH RK MT HH NS AB NK KY TY TK. Contributed reagents/materials/analysis tools: KY MT HH NS AB NK KY TY TK MT. Wrote the paper: KY TK MT.