About the Authors

Manabu Igarashi

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Japan

Kimihito Ito

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Japan

Reiko Yoshida

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Japan

Daisuke Tomabechi

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Japan

Hiroshi Kida

Affiliations Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Japan, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan, OIE Reference Laboratory for Highly Pathogenic Avian Influenza, Sapporo, Japan

Ayato Takada

atakada@czc.hokudai.ac.jp

Affiliation Department of Global Epidemiology, Hokkaido University Research Center for Zoonosis Control, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MI KI HK AT. Analyzed the data: MI RY DT. Wrote the paper: MI AT.