About the Authors

Yuanquan Song

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Mary A. Selak

Affiliation Children's Hospital of Philadelphia Research Institute, Children's Hospital of Philadelphia and University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Corey T. Watson

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Christopher Coutts

Affiliation Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

Paul C. Scherer

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jessica A. Panzer

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Sarah Gibbs

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Marion O. Scott

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Gregory Willer

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Ronald G. Gregg

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Declan W. Ali

Affiliation Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

Michael J. Bennett

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine and Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America

Rita J. Balice-Gordon

rbaliceg@mail.med.upenn.edu

Affiliation Department of Neuroscience, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YS RJBG. Performed the experiments: YS MAS CTW CC PCS JAP SG MOS GW RGG DA MJB. Analyzed the data: YS RJBG. Wrote the paper: YS RJBG.