About the Authors

Sandeep Ravindran

Affiliation Department of Microbiology and Immunology, Stanford University, Stanford, California, United States of America

Melissa B. Lodoen

Affiliation Department of Microbiology and Immunology, Stanford University, Stanford, California, United States of America

Steven H. L. Verhelst

Affiliation Center for Integrative Protein Science Munich, Chemistry of Biopolymers, Technical University Munich, Freising, Germany

Matthew Bogyo

Affiliation Department of Pathology and Microbiology and Immunology, Stanford University, Stanford, California, United States of America

John C. Boothroyd

jboothr@stanford.edu

Affiliation Department of Microbiology and Immunology, Stanford University, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SR JCB. Performed the experiments: SR. Analyzed the data: SR. Contributed reagents/materials/analysis tools: MB JCB. Wrote the paper: SR. Toxofilin-beta-lactamase expressing parasites and their use in assaying rhoptry secretion, used in figure 3: MBL. Organic synthesis of 4-propargyl-oxy-phenacyl bromide (4-PPB), used in figure 5: SHLV.