About the Authors

Hiroko Morimoto

Contributed equally to this work with: Hiroko Morimoto, Mito Kanatsu-Shinohara

Affiliation Department of Molecular Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Mito Kanatsu-Shinohara

Contributed equally to this work with: Hiroko Morimoto, Mito Kanatsu-Shinohara

Affiliation Department of Molecular Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Seiji Takashima

Affiliation Department of Molecular Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Shinichiro Chuma

Affiliation Department of Development and Differentiation, Institute for Frontier Medical Sciences, Kyoto University, Kyoto, Japan

Norio Nakatsuji

Affiliation Department of Development and Differentiation, Institute for Frontier Medical Sciences, Kyoto University, Kyoto, Japan

Masanori Takehashi

Current address: Laboratory of Pathophysiology and Pharmacotherapeutics, Osaka Ohtani University, Tondabayashi, Osaka, Japan

Affiliation Department of Molecular Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Takashi Shinohara

tshinoha@virus.kyoto-u.ac.jp

Affiliations Department of Molecular Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan, Japan Science and Technology Agency, CREST, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TS. Performed the experiments: HM MKS SC TS. Analyzed the data: HM MKS NN TS. Contributed reagents/materials/analysis tools: ST MT TS. Wrote the paper: MKS TS.