About the Authors

Richard J. Bomphrey

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Simon M. Walker

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Graham K. Taylor

graham.taylor@zoo.ox.ac.uk

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RJB GT. Performed the experiments: RJB. Analyzed the data: RJB SMW GT. Contributed reagents/materials/analysis tools: RJB SMW. Wrote the paper: RJB GT.