About the Authors

John R. Lukens

Affiliation Beirne Carter Center for Immunology Research, Department of Microbiology, University of Virginia, Charlottesville, Virginia, United States of America

Joseph S. Dolina

Affiliation Beirne Carter Center for Immunology Research, Department of Microbiology, University of Virginia, Charlottesville, Virginia, United States of America

Taeg S. Kim

Affiliation Beirne Carter Center for Immunology Research, Department of Microbiology, University of Virginia, Charlottesville, Virginia, United States of America

Robert S. Tacke

Affiliation Beirne Carter Center for Immunology Research, Department of Microbiology, University of Virginia, Charlottesville, Virginia, United States of America

Young S. Hahn

ysh5e@virginia.edu

Affiliation Beirne Carter Center for Immunology Research, Department of Microbiology, University of Virginia, Charlottesville, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JRL YSH. Performed the experiments: JRL JSD TSK RST. Analyzed the data: JRL JSD YSH. Wrote the paper: JRL YSH.