About the Authors

Jeffrey S. Hall

jshall@usgs.gov

Affiliation USGS National Wildlife Health Center, Madison, Wisconsin, United States of America

Hon S. Ip

Affiliation USGS National Wildlife Health Center, Madison, Wisconsin, United States of America

J. Christian Franson

Affiliation USGS National Wildlife Health Center, Madison, Wisconsin, United States of America

Carol Meteyer

Affiliation USGS National Wildlife Health Center, Madison, Wisconsin, United States of America

Sean Nashold

Affiliation USGS National Wildlife Health Center, Madison, Wisconsin, United States of America

Joshua L. TeSlaa

Affiliation USGS National Wildlife Health Center, Madison, Wisconsin, United States of America

John French

Affiliation USGS Patuxent Wildlife Research Center, Laurel, Maryland, United States of America

Patrick Redig

Affiliation University of Minnesota School of Veterinary Medicine, Minneapolis, Minnesota, United States of America

Christopher Brand

Affiliation USGS National Wildlife Health Center, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JSH. Performed the experiments: JSH HSI JCF CUM SN JLT. Analyzed the data: JSH HSI JCF CUM SN JLT JF PR CB. Contributed reagents/materials/analysis tools: JSH HSI JCF JF PR CB. Wrote the paper: JSH HSI JCF CUM JF CB.