About the Authors

Shinya Yamamoto

yamamoto@darwin.c.u-tokyo.ac.jp

Affiliation Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Tatyana Humle

Affiliation Wildlife Research Center, Kyoto University, Sakyo-ku, Kyoto, Japan

Masayuki Tanaka

Affiliation Wildlife Research Center, Kyoto University, Sakyo-ku, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SY TH MT. Performed the experiments: SY MT. Analyzed the data: SY. Wrote the paper: SY TH MT.