About the Authors

Philip Rhodes

Affiliation School of Veterinary Medicine and Science, University of Nottingham, Sutton Bonington, United Kingdom

Jim Craigon

Affiliation School of Biosciences, University of Nottingham, Sutton Bonington, United Kingdom

Clint Gray

Affiliation School of Veterinary Medicine and Science, University of Nottingham, Sutton Bonington, United Kingdom

Stuart M. Rhind

Affiliation Macaulay Land Use Research Institute, Craigiebuckler, Aberdeen, United Kingdom

Paul T. Loughna

Affiliation School of Veterinary Medicine and Science, University of Nottingham, Sutton Bonington, United Kingdom

David S. Gardner

david.gardner@nottingham.ac.uk

Affiliation School of Veterinary Medicine and Science, University of Nottingham, Sutton Bonington, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PR JC SR DSG. Performed the experiments: PR CG SR DSG. Analyzed the data: PR JC CG SR DSG. Contributed reagents/materials/analysis tools: JC PL DSG. Wrote the paper: PR PL DSG. Statistical advisor: JC.