About the Authors

Akiko Hirao

Affiliations Laboratory of Physiology and Pharmacology, School of Advanced Science and Engineering, Waseda University, Shinjuku-ku, Tokyo, Japan, Laboratory of Developmental Biology, School of Human Sciences, Waseda University, Saitama, Japan

Yu Tahara

Affiliation Laboratory of Physiology and Pharmacology, School of Advanced Science and Engineering, Waseda University, Shinjuku-ku, Tokyo, Japan

Ichiro Kimura

Affiliation Laboratory of Developmental Biology, School of Human Sciences, Waseda University, Saitama, Japan

Shigenobu Shibata

shibatas@waseda.jp

Affiliation Laboratory of Physiology and Pharmacology, School of Advanced Science and Engineering, Waseda University, Shinjuku-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IK SS. Performed the experiments: AH YT. Analyzed the data: AH YT. Contributed reagents/materials/analysis tools: IK. Wrote the paper: AH SS.