About the Authors

Murray P. Cox

Affiliation ARL Division of Biotechnology, University of Arizona, Tucson, Arizona, United States of America

David A. Morales

Affiliation Department of Mathematics, University of Arizona, Tucson, Arizona, United States of America

August E. Woerner

Affiliation ARL Division of Biotechnology, University of Arizona, Tucson, Arizona, United States of America

Jesse Sozanski

Affiliation ARL Division of Biotechnology, University of Arizona, Tucson, Arizona, United States of America

Jeffrey D. Wall

Affiliation Institute for Human Genetics and Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America

Michael F. Hammer

mfh@u.arizona.edu

Affiliation ARL Division of Biotechnology, University of Arizona, Tucson, Arizona, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JDW MFH. Performed the experiments: AEW. Analyzed the data: MPC DAM JS JDW MFH. Wrote the paper: MPC DAM JDW MFH.