About the Authors

Jennifer Ehren

Contributed equally to this work with: Jennifer Ehren, Belen Morón

Affiliation Department of Chemical Engineering, Stanford University, Stanford, California, United States of America

Belen Morón

Contributed equally to this work with: Jennifer Ehren, Belen Morón

Affiliation Department of Chemistry, Stanford University, Stanford, California, United States of America

Edith Martin

Affiliation Department of Chemistry, Stanford University, Stanford, California, United States of America

Michael T. Bethune

Affiliation Department of Biochemistry, Stanford University, Stanford, California, United States of America

Gary M. Gray

Affiliation Department of Medicine, Stanford University, Stanford, California, United States of America

Chaitan Khosla

khosla@stanford.edu

Affiliations Department of Chemical Engineering, Stanford University, Stanford, California, United States of America, Department of Chemistry, Stanford University, Stanford, California, United States of America, Department of Biochemistry, Stanford University, Stanford, California, United States of America

Competing Interests

C.K. is a Director & Stockholder in Alvine Pharmaceuticals, a company that is developing an oral enzyme drug for celiac disease. None of the other authors have any financial interests to disclose.

Author Contributions

Conceived and designed the experiments: JE BM EM MTB GMG CK. Performed the experiments: JE BM EM. Analyzed the data: JE BM EM MTB GMG CK. Contributed reagents/materials/analysis tools: JE BM EM MTB. Wrote the paper: JE BM MTB GMG CK.