About the Authors

Vernon Reynolds

vreynolds@btopenworld.com

Affiliations Emeritus Professor, School of Anthropology, University of Oxford, Oxford, United Kingdom, Budongo Conservation Field Station, Masindi, Uganda and Faculty of Forestry and Nature Conservation, Makerere University, Kampala, Central Region, Uganda

Andrew W. Lloyd

Affiliation Faculty of Science and Engineering, University of Brighton, Brighton, United Kingdom

Fred Babweteera

Affiliation Budongo Conservation Field Station, Masindi, Uganda and Faculty of Forestry and Nature Conservation, Makerere University, Kampala, Central Region, Uganda

Christopher J. English

Affiliation Faculty of Science and Engineering, University of Brighton, Brighton, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VR. Performed the experiments: FB. Analyzed the data: AL CE. Contributed reagents/materials/analysis tools: AL. Wrote the paper: VR AL FB.