About the Authors

Hui H. Zhang

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Jingxiang Huang

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Katrin Düvel

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Bernard Boback

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Shulin Wu

Affiliation Urologic Research Laboratory, Massachusetts General Hospital, Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Rachel M. Squillace

Affiliation Biology Department, ARIAD Pharmaceuticals, Inc., Cambridge, Massachusetts, United States of America

Chin-Lee Wu

Affiliation Urologic Research Laboratory, Massachusetts General Hospital, Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Brendan D. Manning

bmanning@hsph.harvard.edu

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HHZ JH KD RMS BDM. Performed the experiments: HHZ JH KD BB SW RMS CLW. Analyzed the data: HHZ JH KD RMS CLW BDM. Wrote the paper: HHZ BDM.