About the Authors

Bert van der Zwaag

Contributed equally to this work with: Bert van der Zwaag, Lude Franke

Affiliation Department of Neuroscience and Pharmacology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Lude Franke

Contributed equally to this work with: Bert van der Zwaag, Lude Franke

Affiliations Complex Genetics Section, Department of Biomedical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands, Department of Genetics, University Medical Center Groningen, and University of Groningen, Groningen, the Netherlands

Martin Poot

Affiliation Department of Medical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands

Ron Hochstenbach

Affiliation Department of Medical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands

Henk A. Spierenburg

Affiliation Department of Neuroscience and Pharmacology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Jacob A. S. Vorstman

Affiliation Department of Child and Adolescent Psychiatry, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Emma van Daalen

Affiliation Department of Child and Adolescent Psychiatry, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Maretha V. de Jonge

Affiliation Department of Child and Adolescent Psychiatry, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Nienke E. Verbeek

Affiliation Department of Medical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands

Eva H. Brilstra

Affiliation Department of Medical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands

Ruben van 't Slot

Affiliation Complex Genetics Section, Department of Biomedical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands

Roel A. Ophoff

Affiliation Complex Genetics Section, Department of Biomedical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands

Michael A. van Es

Affiliation Department of Neurology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Hylke M. Blauw

Affiliation Department of Neurology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Jan H. Veldink

Affiliation Department of Neurology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Jacobine E. Buizer-Voskamp

Affiliations Complex Genetics Section, Department of Biomedical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands, Department of Psychiatry, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Frits A. Beemer

Affiliation Department of Medical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands

Leonard H. van den Berg

Affiliation Department of Neurology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Cisca Wijmenga

Affiliations Complex Genetics Section, Department of Biomedical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands, Department of Genetics, University Medical Center Groningen, and University of Groningen, Groningen, the Netherlands

Hans Kristian Ploos van Amstel

Affiliation Department of Medical Genetics, University Medical Center Utrecht, Utrecht, the Netherlands

Herman van Engeland

Affiliation Department of Child and Adolescent Psychiatry, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

J. Peter H. Burbach

J.P.H.Burbach@Umcutrecht.nl (JPHB); W.Staal@Umcutrecht.nl (WGS)

Affiliation Department of Neuroscience and Pharmacology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Wouter G. Staal

J.P.H.Burbach@Umcutrecht.nl (JPHB); W.Staal@Umcutrecht.nl (WGS)

Affiliation Department of Child and Adolescent Psychiatry, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, the Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BvdZ MP RH CW HKPvA HvE JPHB WGS. Performed the experiments: BvdZ LF MP HAS EvD MdJ NV EHB RvS MAVE HMB JV. Analyzed the data: BvdZ LF MP HAS JASV EvD MdJ NV EHB RvS RO MAVE HMB JV JEBV FAB JPHB WGS. Contributed reagents/materials/analysis tools: BvdZ LF MP RH JASV EvD MdJ NV EHB HMB JV FAB LHvdB CW HKPvA HvE JPHB WGS. Wrote the paper: BvdZ LF JPHB WGS.