About the Authors

Wayne D. Crill

wcrill@cdc.gov

Affiliation Arbovirus Diseases Branch, Division of Vector-Borne Infectious Diseases, Centers for Disease Control and Prevention, Public Health Service, U. S. Department of Health and Human Service, Fort Collins, Colorado, United States of America

Holly R. Hughes

Affiliation Arbovirus Diseases Branch, Division of Vector-Borne Infectious Diseases, Centers for Disease Control and Prevention, Public Health Service, U. S. Department of Health and Human Service, Fort Collins, Colorado, United States of America

Mark J. Delorey

Affiliation Arbovirus Diseases Branch, Division of Vector-Borne Infectious Diseases, Centers for Disease Control and Prevention, Public Health Service, U. S. Department of Health and Human Service, Fort Collins, Colorado, United States of America

Gwong-Jen J. Chang

Affiliation Arbovirus Diseases Branch, Division of Vector-Borne Infectious Diseases, Centers for Disease Control and Prevention, Public Health Service, U. S. Department of Health and Human Service, Fort Collins, Colorado, United States of America

Competing Interests

Wayne D. Crill and Gwong-Jen J. Chang are listed as con-inventors for the CDC, US Government on a patent pending for the use of eptitope specific virus-like particles presented in this work as potential serodiagnostic reagents and/or as DNA or subunit vaccines.

Author Contributions

Conceived and designed the experiments: WDC GJJC. Performed the experiments: HRH. Analyzed the data: WDC HRH MD. Contributed reagents/materials/analysis tools: WDC GJJC. Wrote the paper: WDC.