About the Authors

Peter A. Branney

Affiliation Department of Biology, University of York, York, United Kingdom

Laura Faas

Affiliation Department of Biology, University of York, York, United Kingdom

Sarah E. Steane

Affiliation Department of Biology, University of York, York, United Kingdom

Mary Elizabeth Pownall

Affiliation Department of Biology, University of York, York, United Kingdom

Harry V. Isaacs

hvi1@york.ac.uk

Affiliation Department of Biology, University of York, York, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MEp HVI. Performed the experiments: PAB LF SS MEp. Analyzed the data: PAB HVI. Wrote the paper: LF HVI.