About the Authors

Colin R. Lickwar

Contributed equally to this work with: Colin R. Lickwar, Bhargavi Rao

Affiliations Curriculum in Genetics and Molecular Biology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America, Department of Biology, Carolina Center for the Genome Sciences, and Lineberger Comprehensive Cancer Center, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Bhargavi Rao

Contributed equally to this work with: Colin R. Lickwar, Bhargavi Rao

Affiliations Curriculum in Genetics and Molecular Biology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America, Department of Biology, Carolina Center for the Genome Sciences, and Lineberger Comprehensive Cancer Center, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Andrey A. Shabalin

Affiliation Department of Statistics and Operations Research, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Andrew B. Nobel

Affiliation Department of Statistics and Operations Research, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Brian D. Strahl

Affiliations Curriculum in Genetics and Molecular Biology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America, Department of Biochemistry and Biophysics, University of North Carolina School of Medicine, Chapel Hill, North Carolina, United States of America

Jason D. Lieb

jlieb@bio.unc.edu

Affiliations Curriculum in Genetics and Molecular Biology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America, Department of Biology, Carolina Center for the Genome Sciences, and Lineberger Comprehensive Cancer Center, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BS JDL. Performed the experiments: BR. Analyzed the data: CRL AAS AN BS JDL. Wrote the paper: CRL AAS AN BS JDL.