About the Authors

Nicholas J. Haley

Affiliation Department of Microbiology, Immunology, and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Davis M. Seelig

Affiliation Department of Microbiology, Immunology, and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Mark D. Zabel

Affiliation Department of Microbiology, Immunology, and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Glenn C. Telling

Affiliation Department of Molecular Biology and Genetics, University of Kentucky, Lexington, Kentucky, United States of America

Edward A. Hoover

Edward.Hoover@colostate.edu

Affiliation Department of Microbiology, Immunology, and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NJH EAH. Performed the experiments: NJH DMS. Analyzed the data: NJH DMS MDZ GCT EAH. Contributed reagents/materials/analysis tools: NJH MDZ GCT EAH. Wrote the paper: NJH DMS MDZ GCT EAH.