About the Authors

Makoto Hirata

Contributed equally to this work with: Makoto Hirata, Fumitaka Kugimiya

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Fumitaka Kugimiya

Contributed equally to this work with: Makoto Hirata, Fumitaka Kugimiya

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Atsushi Fukai

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Shinsuke Ohba

Affiliation Center for Disease Biology and Integrative Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Naohiro Kawamura

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Toru Ogasawara

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Yosuke Kawasaki

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Taku Saito

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Fumiko Yano

Affiliation Center for Disease Biology and Integrative Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Toshiyuki Ikeda

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Kozo Nakamura

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Ung-il Chung

Affiliation Center for Disease Biology and Integrative Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Hiroshi Kawaguchi

kawaguchi-ort@h.u-tokyo.ac.jp

Affiliation Departments of Sensory & Motor System Medicine, Faculty of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MH FK TI KN UiC HK. Performed the experiments: MH FK AF SO TO. Analyzed the data: MH FK AF. Contributed reagents/materials/analysis tools: SO NK TO YK TS FY TI UiC. Wrote the paper: MH NK HK.