About the Authors

Philip D. Gingerich

gingeric@umich.edu

Affiliation Museum of Paleontology and Department of Geological Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Munir ul-Haq

Affiliation Geological Survey of Pakistan, Quetta, Pakistan

Wighart von Koenigswald

Affiliation Steinmann-Institut für Geologie, Mineralogie und Paläontologie, Universität Bonn, Bonn, Germany

William J. Sanders

Affiliation Museum of Paleontology, University of Michigan, Ann Arbor, Michigan, United States of America

B. Holly Smith

Affiliation Museum of Anthropology, University of Michigan, Ann Arbor, Michigan, United States of America

Iyad S. Zalmout

Affiliation Museum of Paleontology and Department of Geological Sciences, University of Michigan, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Performed the experiments: ISZ. Analyzed the data: PDG MuH WvK WJS BHS. Wrote the paper: PDG MuH WvK WJS BHS ISZ. Participated in field work: PDG MuH ISZ. Contributed geological interpretations: PDG ISZ MuH. Developed chronological framework: PDG MuH ISZ. Carried out functional analysis of skeletons: PDG WJS ISZ. Interpreted life history: PDG BHS.