About the Authors

Susan Broughton

Contributed equally to this work with: Susan Broughton, Nazif Alic

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Nazif Alic

Contributed equally to this work with: Susan Broughton, Nazif Alic

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Cathy Slack

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Timothy Bass

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Tomoatsu Ikeya

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Giovanna Vinti

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Anna Maria Tommasi

Current address: TMRC, James Black Centre, University of Dundee Life Sciences Centre, Dundee, United Kingdom

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Yasmine Driege

Current address: VIB Department for Molecular Biomedical Research, Ghent University, Gent, Belgium

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Ernst Hafen

Affiliation ETH Zurich, Institute of Molecular Systems Biology, Zurich, Switzerland

Linda Partridge

l.partridge@ucl.ac.uk

Affiliation UCL Institute of Healthy Ageing, GEE, University College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SB NA EH LP. Performed the experiments: SB NA CS TB GV AMT YD. Analyzed the data: SB NA CS TB. Contributed reagents/materials/analysis tools: TI EH. Wrote the paper: SB NA LP.