About the Authors

Richard G. Carroll

Contributed equally to this work with: Richard G. Carroll, Carmine Carpenito, Xiaochuan Shan

Affiliations Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Carmine Carpenito

Contributed equally to this work with: Richard G. Carroll, Carmine Carpenito, Xiaochuan Shan

Affiliation Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Xiaochuan Shan

Contributed equally to this work with: Richard G. Carroll, Carmine Carpenito, Xiaochuan Shan

Affiliation Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Gwenn Danet-Desnoyers

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Ronghua Liu

Affiliation Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Shuguang Jiang

Affiliation Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Steven M. Albelda

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Tatiana Golovina

Affiliation Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

George Coukos

Affiliations Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Department of Obstetrics and Gynecology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

James L. Riley

Affiliations Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Zdenka L. Jonak

Affiliation GlaxoSmithKline, Biopharm-CEDD, Biology US, King of Prussia, Pennsylvania, United States of America

Carl H. June

cjune@mail.med.upenn.edu

Affiliations Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

Z.L.J. is an employee of GlaxoSmithKline; the other authors have no competing financial interests to declare.

Author Contributions

Conceived and designed the experiments: RGC SMA GC JLR CHJ. Performed the experiments: RGC CC XS GDD RL TG SJ. Analyzed the data: RGC CC XS GDD TG JLR ZLJ SJ. Contributed reagents/materials/analysis tools: ZLJ. Wrote the paper: RGC CHJ.