About the Authors

Masashi Fujihara

Affiliation Wilmer Eye Institute, Johns Hopkins School of Medicine, Baltimore, Maryland, United States of America

Norihiro Nagai

Affiliation Wilmer Eye Institute, Johns Hopkins School of Medicine, Baltimore, Maryland, United States of America

Thomas E. Sussan

Affiliation Department of Environmental Health Sciences, Bloomberg School of Public Health, Johns Hopkins University, Baltimore, Maryland, United States of America

Shyam Biswal

Affiliation Department of Environmental Health Sciences, Bloomberg School of Public Health, Johns Hopkins University, Baltimore, Maryland, United States of America

James T. Handa

jthanda@jhmi.edu

Affiliation Wilmer Eye Institute, Johns Hopkins School of Medicine, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TES SB JTH. Performed the experiments: MF NN TES. Analyzed the data: MF NN TES SB JTH. Contributed reagents/materials/analysis tools: SB JTH. Wrote the paper: MF SB JTH.