About the Authors

Ching-Shan Chou

Affiliation Department of Mathematics, Center for Mathematical and Computational Biology, Center for Complex Biological Systems, University of California Irvine, Irvine, California, United States of America

Qing Nie

Contributed equally to this work with: Qing Nie, Tau-Mu Yi

Affiliation Department of Mathematics, Center for Mathematical and Computational Biology, Center for Complex Biological Systems, University of California Irvine, Irvine, California, United States of America

Tau-Mu Yi

Contributed equally to this work with: Qing Nie, Tau-Mu Yi

tmy@uci.edu

Affiliation Department of Developmental and Cell Biology, Center for Complex Biological Systems, University of California Irvine, Irvine, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CSC QN TMY. Performed the experiments: CSC. Analyzed the data: CSC QN TMY. Contributed reagents/materials/analysis tools: CSC QN. Wrote the paper: TMY.