About the Authors

Thierry M. Work

thierry_work@usgs.gov

Affiliation U. S. Geological Survey-National Wildlife Health Center, Honolulu Field Station, Honolulu, Hawaii, United States of America

Greta S. Aeby

Affiliation University of Hawaii, Hawaii Institute of Marine Biology, Kaneohe, Hawaii, United States of America

James E. Maragos

Affiliation U. S. Fish & Wildlife Service, Pacific Islands Refuges, Honolulu, Hawaii, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TMW GSA. Performed the experiments: TMW GSA. Analyzed the data: TMW GSA JEM. Contributed reagents/materials/analysis tools: TMW GSA. Wrote the paper: TMW GSA JEM.