About the Authors

Giselle Chamberlain

Affiliation Leopold Muller Arthritis Research Centre, Medical School, Keele University, RJAH Orthopaedic Hospital, Oswestry, Shropshire, United Kingdom

Karina Wright

Affiliation Spinal Studies, Medical School, Keele University, RJAH Orthopaedic Hospital, Oswestry, Shropshire, United Kingdom

Antal Rot

Affiliation Novartis Institutes for BioMedical Research, Vienna, Austria

Brian Ashton

Affiliation Leopold Muller Arthritis Research Centre, Medical School, Keele University, RJAH Orthopaedic Hospital, Oswestry, Shropshire, United Kingdom

Jim Middleton

jim.middleton@rjah.nhs.uk

Affiliation Leopold Muller Arthritis Research Centre, Medical School, Keele University, RJAH Orthopaedic Hospital, Oswestry, Shropshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GC AR BA JFM. Performed the experiments: GC. Analyzed the data: GC. Contributed reagents/materials/analysis tools: KW AR. Wrote the paper: GC JFM.