About the Authors

Josiah Johnston

Affiliation Laboratory of Genetics, NIA Intramural Research Program, National Institutes of Health, Baltimore, Maryland, United States of America

Wendy B. Iser

Affiliation Laboratory of Neurosciences, NIA Intramural Research Program, National Institutes of Health, Baltimore, Maryland, United States of America

David K. Chow

Affiliation Laboratory of Neurosciences, NIA Intramural Research Program, National Institutes of Health, Baltimore, Maryland, United States of America

Ilya G. Goldberg

igg@nih.gov (IG); wolkowca@grc.nia.nih.gov (CW)

Affiliation Laboratory of Genetics, NIA Intramural Research Program, National Institutes of Health, Baltimore, Maryland, United States of America

Catherine A. Wolkow

igg@nih.gov (IG); wolkowca@grc.nia.nih.gov (CW)

Affiliation Laboratory of Neurosciences, NIA Intramural Research Program, National Institutes of Health, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJ IGG CAW. Performed the experiments: JJ WBI DKC CAW. Analyzed the data: JJ IGG CAW. Contributed reagents/materials/analysis tools: JJ WBI DKC IGG. Wrote the paper: JJ IGG CAW.