About the Authors

Tarryne Burke

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, University of KwaZulu-Natal, Durban, South Africa

Bruce Page

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, University of KwaZulu-Natal, Durban, South Africa

Gus Van Dyk

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, University of KwaZulu-Natal, Durban, South Africa

Josh Millspaugh

Affiliation Department of Fisheries and Wildlife Sciences, University of Missouri, Columbia, Missouri, United States of America

Rob Slotow

slotow@ukzn.ac.za

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, University of KwaZulu-Natal, Durban, South Africa

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RS TB GV. Performed the experiments: TB GV. Analyzed the data: RS TB. Contributed reagents/materials/analysis tools: JM. Wrote the paper: RS TB BP JM. Other: Performed the Hormone Assays: JM.