About the Authors

Ada De Luigi

Affiliation Department of Molecular Biochemistry and Pharmacology, Istituto di Ricerche Farmacologiche “Mario Negri”, Milano, Italy

Laura Colombo

Affiliation Department of Molecular Biochemistry and Pharmacology, Istituto di Ricerche Farmacologiche “Mario Negri”, Milano, Italy

Luisa Diomede

Affiliation Department of Molecular Biochemistry and Pharmacology, Istituto di Ricerche Farmacologiche “Mario Negri”, Milano, Italy

Raffaella Capobianco

Affiliation Fondazione I.R.C.C.S. Istituto Neurologico “Carlo Besta”, Milano, Italy

Michela Mangieri

Affiliation Fondazione I.R.C.C.S. Istituto Neurologico “Carlo Besta”, Milano, Italy

Claudia Miccolo

Affiliation Fondazione I.R.C.C.S. Istituto Neurologico “Carlo Besta”, Milano, Italy

Lucia Limido

Affiliation Fondazione I.R.C.C.S. Istituto Neurologico “Carlo Besta”, Milano, Italy

Gianluigi Forloni

Affiliation Department of Neuroscience, Istituto di Ricerche Farmacologiche “Mario Negri”, Milano, Italy

Fabrizio Tagliavini

Affiliation Fondazione I.R.C.C.S. Istituto Neurologico “Carlo Besta”, Milano, Italy

Mario Salmona

salmona@marionegri.it

Affiliation Department of Molecular Biochemistry and Pharmacology, Istituto di Ricerche Farmacologiche “Mario Negri”, Milano, Italy

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FT RC GF MS AD LC. Performed the experiments: RC MM CM AD LC LD LL. Analyzed the data: FT RC MM CM MS AD LC LD LL. Wrote the paper: MS AD.