About the Authors

Dick S. J. Groenenberg

groenenberg@naturalis.nl

Affiliation National Museum of Natural History Naturalis, Leiden, The Netherlands

Albert J. Beintema

Affiliation Biowrite, Gorssel, The Netherlands

René W. R. J. Dekker

Affiliation National Museum of Natural History Naturalis, Leiden, The Netherlands

Edmund Gittenberger

Affiliations National Museum of Natural History Naturalis, Leiden, The Netherlands, Institute of Biology, Leiden University, Leiden, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DG. Performed the experiments: DG. Analyzed the data: DG. Wrote the paper: DG. Other: Provided a hypothesis and some of the specimens: AB. Reviewed the manuscript: EG RD AB.