About the Authors

Amr H. Sawalha

*E-mail: amr-sawalha@omrf.ouhsc.edu

Affiliations Department of Medicine, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America, United States Department of Veterans Affairs Medical Center, Oklahoma City, Oklahoma, United States of America, Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Ryan Webb

Affiliation Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Shizhong Han

Affiliation Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Jennifer A. Kelly

Affiliation Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Kenneth M. Kaufman

Affiliations Department of Medicine, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America, United States Department of Veterans Affairs Medical Center, Oklahoma City, Oklahoma, United States of America, Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Robert P. Kimberly

Affiliation Division of Clinical Immunology and Rheumatology, Department of Medicine, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Marta E. Alarcón-Riquelme

Affiliation Department of Genetics and Pathology, Uppsala University, Uppsala, Sweden

Judith A. James

Affiliations Department of Medicine, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America, Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Timothy J. Vyse

Affiliation Rheumatology Section, Imperial College, Hammersmith Hospital, London, United Kingdom

Gary S. Gilkeson

Affiliation Division of Rheumatology, Department of Medicine, Medical University of South Carolina, Charleston, South Carolina, United States of America

Chan-Bum Choi

Affiliation Division of Rheumatology, Department of Internal Medicine and the Hospital for Rheumatic Diseases, Hanyang University, Seoul, Republic of Korea

R. Hal Scofield

Affiliations Department of Medicine, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America, United States Department of Veterans Affairs Medical Center, Oklahoma City, Oklahoma, United States of America, Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Sang-Cheol Bae

Affiliation Division of Rheumatology, Department of Internal Medicine and the Hospital for Rheumatic Diseases, Hanyang University, Seoul, Republic of Korea

Swapan K. Nath

Affiliation Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

John B. Harley

Affiliations Department of Medicine, University of Oklahoma Health Sciences Center, Oklahoma City, Oklahoma, United States of America, United States Department of Veterans Affairs Medical Center, Oklahoma City, Oklahoma, United States of America, Arthritis and Immunology Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JH AS KK. Performed the experiments: RW KK. Analyzed the data: AS SN SH JK. Contributed reagents/materials/analysis tools: GG JH JJ RK RS TV SB MA CC. Wrote the paper: AS.