About the Authors

Koichi Iijima

*E-mail: Koichi.Iijima@jefferson.edu (KI); zhongyi@cshl.edu (YZ)

Affiliations Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America, Laboratory of Neurodegenerative Diseases and Gene Discovery, Farber Institute for Neurosciences, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Department of Biochemistry and Molecular Biology, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Hsueh-Cheng Chiang

Affiliations Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America, Department of Neurobiology and Behavior, State University of New York, Stony Brook, New York, United States of America

Stephen A. Hearn

Affiliation Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America

Inessa Hakker

Affiliation Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America

Anthony Gatt

Affiliation Laboratory of Neurodegenerative Diseases and Gene Discovery, Farber Institute for Neurosciences, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Christopher Shenton

Affiliation Laboratory of Neurogenetics and Protein Misfolding Diseases, Farber Institute for Neurosciences, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Linda Granger

Affiliations Laboratory of Neurodegenerative Diseases and Gene Discovery, Farber Institute for Neurosciences, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Laboratory of Neurogenetics and Protein Misfolding Diseases, Farber Institute for Neurosciences, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Amy Leung

Affiliation Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America

Kanae Iijima-Ando

Affiliations Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America, Laboratory of Neurogenetics and Protein Misfolding Diseases, Farber Institute for Neurosciences, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Department of Biochemistry and Molecular Biology, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Yi Zhong

*E-mail: Koichi.Iijima@jefferson.edu (KI); zhongyi@cshl.edu (YZ)

Affiliation Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KI KI-A. Performed the experiments: IH KI HC SH AL KI-A AG LG CS. Analyzed the data: YZ KI HC SH KI-A. Contributed reagents/materials/analysis tools: KI. Wrote the paper: YZ KI KI-A.