About the Authors

Richard A. Holland

*E-mail: rahollan@Princeton.edu

Affiliations Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America, Institute of Integrative and Comparative Biology, University of Leeds, Leeds, United Kingdom

Joseph L. Kirschvink

Affiliation Division of Geological and Planetary Sciences, California Institute of Technology, Pasadena, California, United States of America

Thomas G. Doak

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Martin Wikelski

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MW RH JK. Performed the experiments: MW RH TD. Analyzed the data: RH. Contributed reagents/materials/analysis tools: MW RH JK. Wrote the paper: MW RH JK TD.