About the Authors

Timothy J. Page

*E-mail: t.page@griffith.edu.au

Affiliation Australian Rivers Institute, Griffith University, Nathan, Queensland, Australia

William F. Humphreys

Affiliation Western Australian Museum, Welshpool DC, Western Australia, Australia

Jane M. Hughes

Affiliation Australian Rivers Institute, Griffith University, Nathan, Queensland, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JH TP WH. Performed the experiments: TP. Analyzed the data: TP. Contributed reagents/materials/analysis tools: TP WH. Wrote the paper: JH TP WH.