About the Authors

Toby G. Knowles

To whom correspondence should be addressed. E-mail: Toby.Knowles@bristol.ac.uk

Affiliation School of Veterinary Science, University of Bristol, Langford, Bristol, United Kingdom

Steve C. Kestin

Affiliation School of Veterinary Science, University of Bristol, Langford, Bristol, United Kingdom

Susan M. Haslam

Affiliation School of Veterinary Science, University of Bristol, Langford, Bristol, United Kingdom

Steven N. Brown

Affiliation School of Veterinary Science, University of Bristol, Langford, Bristol, United Kingdom

Laura E. Green

Affiliation Ecology and Epidemiology Group, Biological Sciences, University of Warwick, Coventry, United Kingdom

Andrew Butterworth

Affiliation School of Veterinary Science, University of Bristol, Langford, Bristol, United Kingdom

Stuart J. Pope

Affiliation School of Veterinary Science, University of Bristol, Langford, Bristol, United Kingdom

Dirk Pfeiffer

Affiliation The Royal Veterinary College, Royal College Street, London, United Kingdom

Christine J. Nicol

Affiliation School of Veterinary Science, University of Bristol, Langford, Bristol, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DP CN TK SK SB AB LG. Performed the experiments: TK SK SH SB AB SP. Analyzed the data: DP CN TK SK SH SB LG. Wrote the paper: DP CN TK SK SH LG.