About the Authors

Tadashi Iwami

Contributed equally to this work with: Tadashi Iwami, Osamu Kishida, Kinya Nishimura

Affiliation Graduate School of Fisheries Sciences, Hokkaido University, Hakodate, Hokkaido, Japan

Osamu Kishida

Contributed equally to this work with: Tadashi Iwami, Osamu Kishida, Kinya Nishimura

Current address: Center for Ecological Research, Kyoto University, Otsu, Shiga, Japan

Affiliation Graduate School of Fisheries Sciences, Hokkaido University, Hakodate, Hokkaido, Japan

Kinya Nishimura

Contributed equally to this work with: Tadashi Iwami, Osamu Kishida, Kinya Nishimura

To whom correspondence should be addressed. E-mail: kinya@fish.hokudai.ac.jp

Affiliation Graduate School of Fisheries Sciences, Hokkaido University, Hakodate, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: OK KN TI. Performed the experiments: OK TI. Analyzed the data: OK TI. Contributed reagents/materials/analysis tools: OK TI. Wrote the paper: OK KN TI. Other: Designed the research program: TI OK KN. Conducted field research and laboratory experiments: TI OK. Discussed the results: TI OK KN. Commented on the manuscript: TI OK KN.