About the Authors

Naohiro Kawamura

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Fumitaka Kugimiya

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Yasushi Oshima

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Shinsuke Ohba

Affiliation Center for Disease Biology and Integrative Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Toshiyuki Ikeda

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Taku Saito

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Yusuke Shinoda

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Yosuke Kawasaki

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Naoshi Ogata

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Kazuto Hoshi

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Toru Akiyama

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

William S. Chen

Affiliation Transgenic and Knockout Mouse Laboratory, University of Kansas, Lawrence, Lawrence, Kansas, United States of America

Nissim Hay

Affiliation Department of Molecular Genetics, College of Medicine, University of Illinois at Chicago, Chicago, Illinois, United States of America

Kazuyuki Tobe

Affiliation Department of Metabolic Diseases, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Takashi Kadowaki

Affiliation Department of Metabolic Diseases, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Yoshiaki Azuma

Affiliation Teijin Institute for Biomedical Research, Tokyo, Japan

Sakae Tanaka

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Kozo Nakamura

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Ung-il Chung

Affiliation Center for Disease Biology and Integrative Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Hiroshi Kawaguchi

To whom correspondence should be addressed. E-mail: kawaguchi-ort@h.u-tokyo.ac.jp

Affiliation Department of Sensory and Motor System Medicine, Faculty of Medicine, University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HK NK FK TA KN UC. Performed the experiments: NK FK YO SO TI TS YK YS YA. Analyzed the data: HK NK FK TI NO KH TA KN UC ST. Contributed reagents/materials/analysis tools: NH TK YO KT ST WC. Wrote the paper: HK NK.