About the Authors

Michal Grzmil

Affiliation Sir William Dunn School of Pathology, University of Oxford, Oxford, United Kingdom

Danny Whiting

Affiliation Sir William Dunn School of Pathology, University of Oxford, Oxford, United Kingdom

John Maule

Affiliation MRC Human Genetics Unit and University of Edinburgh Cancer Research Centre, Western General Hospital, Edinburgh, United Kingdom

Corina Anastasaki

Affiliation MRC Human Genetics Unit and University of Edinburgh Cancer Research Centre, Western General Hospital, Edinburgh, United Kingdom

James F. Amatruda

Affiliation Departments of Pediatrics, Internal Medicine and Molecular Biology, University of Texas Southwestern Medical Center, Dallas, Texas, United States of America

Robert N. Kelsh

Affiliation Centre for Regenerative Medicine, Developmental Biology Programme, Department of Biology and Biochemistry, University of Bath, Bath, United Kingdom

Chris J. Norbury

Affiliation Sir William Dunn School of Pathology, University of Oxford, Oxford, United Kingdom

E. Elizabeth Patton

To whom correspondence should be addressed. E-mail: epatton@staffmail.ed.ac.uk

Affiliations Sir William Dunn School of Pathology, University of Oxford, Oxford, United Kingdom, MRC Human Genetics Unit and University of Edinburgh Cancer Research Centre, Western General Hospital, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EP CN MG. Performed the experiments: JA EP JM CA MG DW. Analyzed the data: JA EP CN. Contributed reagents/materials/analysis tools: RK EP CN. Wrote the paper: EP.