About the Authors

Yoshihito Niimura

To whom correspondence should be addressed. E-mail: niimura@bioinfo.tmd.ac.jp

Affiliations Department of Bioinformatics, Medical Research Institute, Tokyo Medical and Dental University, Tokyo, Japan, Institute of Molecular Evolutionary Genetics and Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Masatoshi Nei

Affiliation Institute of Molecular Evolutionary Genetics and Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MN YN. Performed the experiments: YN. Analyzed the data: YN. Contributed reagents/materials/analysis tools: YN. Wrote the paper: MN YN.